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Tuesday, July 1, 2014

A Passel of Pollinators

The third week in June was National Pollinator Week.  Our own GTM Research Reserve didn't celebrate pollinators for just one week.  This author wrote a batch of "Weird Animal Wednesday" articles for their Facebook page about ALL the critters that pollinate our local flora.  Here are links to the passel.  And in case you don't know what a "passel" means, it means a bunch, a lot, a collection of indeterminate number... (Click the Read More Link below.)



On May 14, the pink-striped hawkmoth (also known as the sphinxmoth) got a shout-out, but it looks like it got lost somewhere on Facebook.  Well, here it is for your reading pleasure:


Off to the (Moth) Races

Each spring my faithful brugmansia suaveolens (angel’s trumpet) breaks the soil, green stalk spewing leaves as it races skyward.  By June it is six feet tall and covered in a blush of drooping flowers that unfurl to release their fragrance only at night.  Why the pagentry only when darkness comes?  Considering the brugmansia is toxic, as are the datura (jimsonweed) and belladonna (deadly nightshade) in the Solanaceae family, what shadowy creature would nectar on its flowers?  Could a visitor actually retrieve the juice from the base of the brugmansia’s twelve-inch throat?  

Enter the Pink winged hawkmoth, agrius cingulata.  This member of the Sphingidae family is as lovely and exotic-looking as the brugmansia flower it visits.  There are some beautiful moths, but the hawkmoth’s sleek wings and tapered body exhibit a geometry that is both stunning and functional.  The name, “hawkmoth”, refers to its incredible flying capability, achieving speeds up to 12 MPH.  Not only are these moths fast, but some are terrific acrobats, able to hover and flit side to side.  The hummingbird hawkmoth is so called because its wings hum. It can hover and dart like a hummer and it looks like one with a two-inch wingspan.  Its habit of daytime feeding causes it to often be mistaken for the tiny bird.  

But back to a. cingulata, the Pink-winged hawkmoth.  Did I mention that it’s pink?  Pink and brownish-gray stripes alternate along its powerful thorax, and blushes of pink hue the lower wings.  With a wingspan of 3 ¾” to 4 ¾” and a body like an oversized Smith & Wesson Special cartridge, it is a sight to see one hovering under an angel’s trumpet.  Not that I have.  But good (or lucky) photographers have caught agrius cingulata feeding in mid-flight.  And those pictures reveal its most impressive feature: the four-inch proboscis.  Uncoiled, this feeding tube is two times longer than its body.  This adaptation, along with its hovering ability, gives the hawkmoth access to the brugmansia’s nectar. 
Hawkmoths, known as sphinxmoths, are nectar experts that specialize in species with elongated or elaborate flowers, including angel’s trumpet, jimsonweed, moonflowers, morning glories, and orchids.  The main character in the book, “The Orchid Thief” and movie, “Adaptation,” tells us that the elusive ghost orchid can only be pollinated by the Giant sphinxmoth, which has a nine-inch proboscis.  Darwin rightly predicted if there’s a flower, no matter the shape, there’s a pollinator adapted to it.  

No matter the toxicity either.  Poisonous trumpet flowers, whose extracts yield belladonna and other opiods have no effect on these hawkmoths.  Mysterious and alluring, one can understand why trumpet flowers and their symbiont lepidoptera were often depicted in the arts.  A favorite Victorian theme visualized Shakespeare’s fairy queen from A Midsummer Night’s Dream, Titania, sleeping under a bower of trumpet flowers while her attendants, Cobweb, Mustardseed, and Moth frolic nearby.  The hawkmoth has also found its way into the modern imagination.  The creature sealing Jody Foster’s lips in the poster for “The Silence of the Lambs” is a stylized Death’s-head hawkmoth.  The murderer buries it’s chrysalis in his victims’ throats.

To understand why it is also called a sphinxmoth, let’s examine hawkmoth’s “salad days” when it was an ungainly, insanely hungry caterpillar.  The larva has a resting pose that reminded biologists of the Egyptian sphinx:  forelegs extended and head tucked in.  Their nickname is “hornworm.”  Agrius cingulata spends its youth as a Sweetpotato hornworm.  A voracious pest, it gorges on sweet potato leaves, morphing from white to  green to brown.  Its mighty anal horn (helpful in misdirecting predator attacks) gradually softens and shrinks prior to pupation.  Southern farmers have a fulltime job keeping hornworms from defoliating potatoes, tomatoes, and other crops.  Oddly, there have been no a. cingulara citings of its imago St. Johns County.  Our farms produce large quantities of its host plant.

Who could foresee agrius cingulata becoming handsome, a Chuck Yeager of the night flyers?  The chrysalis hints of its future beauty:  a furrowed amber cocoon capped by a swirl, it resembles a clasped pendant.  The “clasp” is actually the case for the hawkmoth’s proboscis.  A Pink-striped hawkmoth emerges from its pupa.

But not in my garden.  My brugmansia blossoms beckon nightly.  The unsatisfied blooms soon drop to the ground.  If it has a nectar-loving visitor, it leaves the plant as barren as before.  Adaptation enabled the hawkmoth to reach its prize and survive the feast; it also gave it the “right stuff” to pollinate brugmansia suavolens:  the proper length and shape to perturb its stamen and transport the pollen to a nearby pistil.  In other locales.  But not here. 
I wonder if agrius. cingulata’s absence is fairy mischief.  Perhaps Titania’s heir, inspired by our equestrian Triple Crown, has summoned her First Coast subjects elsewhere:  to attend her nocturnal races.  The steed of choice would surely be the hawkmoth, and I imagine one lucky fairy jockeying in colors of pink and gray.  A stellar flyer, it’d be a wings-down favorite.  If it found itself racing neck-and-neck in the home stretch, a. cingulata could magically win by a proboscis.  


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